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Michael Caldwell and Halle P. Street

Paleontology: Ancient Marine Reptiles is a four-lesson course teaching a comprehensive overview of the evolutionary changes that occur when air-breathing terrestrial animals return to water. This course examines the diversity, adaptations, convergence, and phylogenetic relationships of extinct marine reptiles. Students will explore three major groups of marine reptiles: ichthyosaurs, plesiosaurs, and mosasaurs. Watch a preview of the course here: https://uofa.ualberta.ca/courses/paleontology-marine-reptiles

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Syllabus

Introduction to Marine Reptiles
Welcome to the first lesson of Palaeontology: Ancient Marine Reptiles. In this lesson we will explore the main theme of the course: the aquatic problem. In other words, what happens when a terrestrial animal returns to the water permanently? How do air-breathing, land-lubbing creatures once again adapt to life in the sea? Life in water is very different than life on land. Water is much denser than air, which affects all aspects of an animal's life including movement, sight, and hearing. In addition, animals that return to the water cannot breathe water, and so must return to the surface for air. Water also conducts heat much better than air, making staying warm and active a challenge. Despite all these obstacles, many land animals have returned to the water throughout the course of evolutionary history. In fact, many examples of them are living today including whales, seals, crocodiles, sea turtles and penguins. Each of these animals had ancestors that returned to the water. The process of overcoming the challenges associated with this transition is what we refer to as the aquatic problem. This lesson will explore many different types of adaptations that modern and extinct animals have evolved to meet these challenges. You will be introduced to some extinct groups of reptiles you have probably never heard of, and will gain a new appreciation for how well suited modern marine animals are to their environment. Just a quick note before you get started: 'Palaios' is the Greek word for 'ancient', so palaeontology or paleontology is the study of ancient life. Both spellings are correct, with palaeontology used in Britain, and paleontology more common in the US.
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Ichthyopterygians
Welcome back! We hope that you enjoyed the first lesson of the Ancient Marine Reptiles mini MOOC. In lesson one, we discussed a variety of modern and extinct groups of aquatic amniotes. Some of these groups would have been familiar to you, and some you had probably never heard of before. We also explored some of the many differences between living in water and living on land, and we gave examples of how some amniote groups have overcome these challenging differences. As you may have already seen, some of the solutions to the different aspects of the aquatic problem were solved in the same way by unrelated groups- a phenomenon called convergence. The next three lessons will build on your understanding of the aquatic problem as we investigate it more detail in three extinct groups of marine reptiles: the ichthyopterygians, sauropterygians, and mosasauroids. Each lesson will follow the same general outline. First, we will introduce you to the group and some of its diversity. The second part of each lesson will focus on how that group adapted to solve the aquatic problem, and how this resulted in specializations for feeding, locomotion and reproduction. Finally we will give you an overview of the evolutionary history of the group through time and space, as well as presenting some important fossils and localities from Canada and around the world. In lesson two, we will study the first of the three major groups of extinct marine amniotes: the ichthyopterygians. The ichthyopterygians were highly specialized animals. Most of the later members looked something like a dolphin or a tuna, a great example of convergent evolution. These reptiles were probably among the fastest aquatic creatures that ever lived. They were found all over the world, from the early Triassic to the mid-Cretaceous, a span of 150 million years. By the end of this lesson, you should be familiar with the history and diversity of this group, and be able to appreciate some of their impressive specializations to solve the aquatic problem.
Sauropterygians
Welcome back! We hope you enjoyed the lesson on ichthyopterygians, the first of three in depth explorations of an extinct marine reptile group that we will cover. In this lesson you learned that ichthyosaurs are among the most specialized reptiles that ever lived. They had many adaptations to solving the aquatic problem including large eyes for seeing in deep water, powerful tails to power their thunniform swimming, and two sets of flippers for stability and steering. We also discussed the hypothesis that ichthyosaurs may have overcome the problem of being cold-blooded reptiles living in water by evolving some endothermic capabilities. Even though ichthyopterygians were well adapted for a life in the water, this lineage still went extinct long before the End-Cretaceous Mass Extinction for unknown reasons. The lesson you are about to start will follow a similar format to lesson 2, but instead of ichthyopterygians, we will be investigating a second major marine reptile lineage: the sauropterygians. These animals shared the seas with the ichthyosaurs for much of their 180 million year existence. You may already know some of the members of this group such as the long-necked elasmosaurs and the massive-jawed pliosaurs such as Liopleurodon. In this lesson we will expand your knowledge of these animals by introducing you to their lizard-like ancestors and a wider variety of the derived members. We will present you with one of the biggest unsolved mysteries surrounding marine reptiles: how did plesiosaurs use their four massive, wing-like flippers to swim? We will also look at how they ate and reproduced, and we will finish by taking you on a worldwide tour of plesiosaur diversity through time. We hope you enjoy learning about this group of reptiles, who had adaptations so unique that they have never been seen in any other lineage of marine tetrapods.
Mosasauroids
Welcome to the final lesson in Palaeontology: Ancient Marine Reptiles. The lesson you just finished gave you an in depth look at one of the most morphologically disparate groups of marine tetrapods to ever live. There were the turtle-shaped placodonts with crushing teeth; the extraordinarily long-neck elasmosaurs; and the pliosaurs with jaws massive enough to take on nearly any prey. Even though sauropterygians evolved many different body plans, they all shared certain adaptations for solving the aquatic problem such as live birth and appendicular locomotion. In contrast, the final group we will investigate showed convergence on one morphotype. The mosasaurs, though diverse, were all fairly long reptiles, with large jaws, four flippers and a long, lobed tail. You may recognise one of them from Jurassic World, where an enormous genetically-engineered Mosasaurus leaped out of the water to eat a shark. This last lesson will once again start by examining the diversity of the group, from their terrestrial origins to last days of the Cretaceous when they ruled as apex predators. We will investigate the specific adaptations of the group to the aquatic problem, and finish with an overview of the history of mosasaurs throughout time and space. Please enjoy the last lesson of Palaeontology: Ancient Marine Reptiles. It is an appropriate way to finish this course since the mosasaurs were among the largest and most powerful marine predators to ever live, and were the last major marine reptile group to evolve during the Mesozoic.

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Know what's good
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Explores the evolutionary changes when air-breathing terrestrial animals return to water, which is highly relevant to evolutionary biology
Taught by Michael Caldwell and Halle P. Street, who are recognized for their work in evolutionary biology
Examines the diversity, adaptations, convergence, and phylogenetic relationships of extinct marine reptiles
Offers hands-on interactives, such as a preview of the first lesson
May be suitable for students with some background knowledge, such as introductory science and biology

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Reviews summary

Ancient marine reptiles: paleontology

Learners say this is a largely positive course that provides a thorough overview of three major lineages of marine reptiles that lived during the Mesozoic era: Ichthyosaurs, Plesiosaurs, and Mosasaurs. Based on 192 reviews, students remark that the course is well received, engaging, and informative but may be difficult for those without prior knowledge in paleontology. Notably, students appreciate the enthusiastic instructor, detailed notes, engaging assignments, and well-organized format. The University of Alberta's high-quality production of the course, including maps, diagrams, and artistic recreations, is also widely praised.
The instructor, W. Scott Persons, is very knowledgeable and genuinely passionate about the subject.
"I really enjoyed the course & finding out about the creatures that have fascinated me for years. The course was very well structured in logical sections that enabled me to study when I wanted."
"This course is very good: the presentations were very well done, the drawn images were very unique and stylish, and the information was very interesting. Very well done, definitetly recomend."
"I strongly recommend this course! The narrator, Mr W Scott Persons, was very good at narrating, so enthusiastic and nice. Very informative and this course made me decide to keep on studying Paleontology."
The quizzes and discussion questions are a great way to test your understanding of the material.
"Excellent, very comprehensive course with outstanding supplemental materials. Dr. Persons is an excellent presenter."
"I like to see when the teacher is passionate about his area and knows how to explain it in a didactic and precise way."
"This was a very excellent course as I want to learn more about animals in preparation the study as a veterinary technician. "
The course notes are well-written and provide a comprehensive overview of the material.
"Excellent and very interesting course! The instructor is very good and passionate, highly recommend if you have any interest in Palaeontology. The first week requires a bit of background knowledge, but thankfully the same university offers a Dino 101 course, which the first week of that gave me enough knowledge to continue with this course."
"I have only words of praise for University of Alberta courses. The artistic level is amazing, the notes are detailed and wonderful - instead of just plain transcripts you get a small encyclopaedia! The lectures are fascinating, and you have a chance to see real-life fossils too."
"I really enjoyed the course and finding out about the creatures that have fascinated me for years. The course was very well structured in logical sections that enabled me to study when I wanted."
Some of the information presented in the course is outdated.
"While the information presented in the course is top-notch, it's held back by the use of Flash for many interactive aspects of the videos and also large PDFs for the reading material."
"I thought this was a nice course. Sometimes the questions gave me pause and forced me to think. One complaint I have is that occasionally statements about functional morphology lack nuance."
"This course also seems to have a few bits of outdated info. For example, it is now known that Atopodentatus did not have a “zipper mouth”; this was based on a specimen that had a badly crushed skull."
The material is challenging, but the instructor does a good job of explaining complex concepts.
"The course is great. I have learn a lot. "
"Learned a lot, enthusiastic instructors. "
"F​ascinating material, clearly explained."

Activities

Be better prepared before your course. Deepen your understanding during and after it. Supplement your coursework and achieve mastery of the topics covered in Paleontology: Ancient Marine Reptiles with these activities:
Review previous coursework or study materials on marine biology and paleontology
Refresh knowledge of marine biology and paleontology to strengthen foundation for course
Browse courses on Marine Biology
Show steps
  • Review notes and textbooks from previous courses
  • Complete practice questions or exercises
Review biology and ecology of Reptilia
Review the biology and ecology of Reptilia to help prepare for the main course content
Browse courses on Biology
Show steps
  • Review online materials on Reptilia
  • Read textbook chapters on the biology of Reptilia
  • Practice identifying reptiles in online databases
Practice naming fossil groups and identifying characteristics
Complete practice drills to sharpen fossil identification and naming skills
Browse courses on Paleontology
Show steps
  • Use online resources for fossil identification practice
  • Review textbook sections on fossil identification
Four other activities
Expand to see all activities and additional details
Show all seven activities
Follow video tutorials on marine reptile evolutionary adaptations
Enhance understanding by following video tutorials on marine reptile evolutionary adaptations
Browse courses on Paleontology
Show steps
  • Search for video tutorials on marine reptile adaptations
  • Watch and take notes on the tutorials
  • Apply the learned concepts to course discussions
Participate in online forums to discuss course content
Engage with peers in online forums to clarify concepts and deepen understanding
Browse courses on Paleontology
Show steps
  • Join online forums related to the course
  • Post questions and participate in discussions
Review 'Marine Reptiles' by Michael J. Benton
Expand knowledge of marine reptiles by reviewing 'Marine Reptiles' by Michael J. Benton
Show steps
  • Read selected chapters from 'Marine Reptiles'
  • Summarize key concepts and findings from the book
Create a presentation on a specific group of marine reptiles
Solidify knowledge and develop presentation skills by creating a presentation on a specific group of marine reptiles
Show steps
  • Choose a specific group of marine reptiles to research
  • Gather information on the group's diversity, adaptations, and evolutionary history
  • Create a presentation using slides or other visual aids
  • Present the presentation to the class or online

Career center

Learners who complete Paleontology: Ancient Marine Reptiles will develop knowledge and skills that may be useful to these careers:
Paleontologist
A Paleontologist may explore a variety of extinct species to investigate how different species evolved. This course focuses on ancient marine reptiles. By understanding how they evolved, you may learn how species evolve and adapt over time to fit different environments. This course's introduction to extinct marine reptiles is particularly relevant to a Paleontologist.
Curator
A Curator may work in a museum setting where they would be responsible for preserving historical objects. This course, with its focus on paleontology, explores the history of marine reptiles who lived millions of years ago. The course also touches on some of the fossils and localities where they existed. This knowledge is particularly relevant to a Curator.
Geologist
A Geologist may use scientific methods to explore the physical structure of the earth. In this course on ancient marine reptiles, you will learn about the physical changes that occurred in these species over millions of years and the impact that had on their existence. This course also touches on different localities where these species existed. This course may help build a foundation for understanding how the earth changed over the course of time to allow for human existence.
Marine Biologist
A Marine Biologist uses scientific methods to explore living organisms and their habitats in the ocean. This course on marine reptiles, though extinct, focuses on the evolutionary changes that occur when air-breathing, land-lubbing creatures return to the water. This knowledge may help build a foundation for understanding how marine species evolved and the challenges they faced when adapting to life in the water.
Archaeologist
An Archaeologist may collect and examine fossils, artifacts, and other resources to better understand history. This course, focusing on ancient marine reptiles, may help build a foundation for understanding how archeological methods can be used to uncover fossils and artifacts of ancient species to learn more about our history.
Environmental Scientist
An Environmental Scientist studies the environment and looks for ways to protect it. This course focuses on exploring the evolutionary changes that occur when air-breathing, land-lubbing creatures return to the water. Understanding how species adapt and change to overcome challenges may help you build a foundation for developing ways to protect the earth and its inhabitants.
Science Teacher
A Science Teacher develops lesson plans and teaches science courses. This course focuses on ancient marine reptiles and may help build a foundation for teaching life science courses on evolution and adaptation.
Biologist
A Biologist studies living organisms and their habitats. This course focuses on ancient marine reptiles, however, this course may help build a foundation for understanding how different organisms evolved and adapted to different environments.
Zoologist
A Zoologist studies the behavior, physiology, anatomy, and classification of animals. This course focuses on marine reptiles. By learning about the diversity and adaptations of extinct marine reptiles, you may build a foundation for understanding the diversity and adaptations of modern animals.
Technical Writer
A Technical Writer writes instruction manuals, technical reports, and other technical materials. This course focuses on the evolutionary changes that occur when air-breathing, land-lubbing creatures return to the water. If you were interested in writing about scientific discoveries or prehistoric species, this course may help build a foundation.
Writer
A Writer develops and writes content. This course explores the evolutionary changes that occur when air-breathing, land-lubbing creatures return to the water. If you were interested in writing about scientific discoveries or prehistoric species, this course may help build a foundation.
Journalist
A Journalist trains to gather, analyze, and report on news and current events. This course focuses on the evolutionary changes that occur when air-breathing, land-lubbing creatures return to the water. If you were interested in reporting on scientific discoveries or prehistoric species, this course may help build a foundation.
Science Writer
A Science Writer writes about scientific topics for a general audience. This course explores the evolutionary changes that occur when air-breathing, land-lubbing creatures return to the water. This knowledge may help build a foundation for writing about scientific discoveries or prehistoric species.
Researcher
A Researcher conducts scientific research and experiments. This course focuses on marine reptiles. By exploring the evolutionary changes that occur when air-breathing, land-lubbing creatures return to the water, this course may help build a foundation for conducting research on modern aquatic species.
Editor
An Editor reviews, corrects, and revises written material. This course focuses on the evolutionary changes that occur when air-breathing, land-lubbing creatures return to the water. If you were interested in working with writers who produce scientific discoveries or prehistoric species, this course may help build a foundation.

Reading list

We've selected eight books that we think will supplement your learning. Use these to develop background knowledge, enrich your coursework, and gain a deeper understanding of the topics covered in Paleontology: Ancient Marine Reptiles.
Provides a detailed overview of the evolutionary history of marine reptiles, from their origins in the Permian period to their extinction at the end of the Cretaceous period. It would be a valuable resource for students who want to learn more about the diversity and adaptations of marine reptiles.
Classic work on the functional morphology of vertebrates, including marine reptiles. It provides a detailed overview of the different ways that vertebrates have adapted to their environment and how their morphology reflects their function. It would be a valuable resource for students who want to learn more about the adaptations of marine reptiles.
Provides a detailed overview of the origin and evolution of marine reptiles. It would be a valuable resource for students who want to learn more about the history of marine reptiles and their relationships to other groups of reptiles.
Provides a detailed overview of the paleobiology of marine reptiles, including their ecology, behavior, and extinction. It would be a valuable resource for students who want to learn more about the history of marine reptiles and their role in marine ecosystems.
Provides a detailed overview of the ecology of marine reptiles, including their habitat, behavior, and feeding habits. It would be a valuable resource for students who want to learn more about the biology of marine reptiles and their role in marine ecosystems.
Provides a comprehensive overview of vertebrate biology, including a section on marine reptiles. It would be a useful reference for students who want to learn more about the biology of marine reptiles and their role in marine ecosystems.
Provides a comprehensive overview of vertebrate paleontology and evolution, including a section on marine reptiles. It would be a useful reference for students who want to learn more about the history of marine reptiles and their role in marine ecosystems.
Provides a broad overview of vertebrate paleontology and evolution, covering the major groups of vertebrates from fish to mammals. It would be a useful reference for students who want to learn more about the evolutionary history of marine reptiles.

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